Google
has been in the search game a long time, and saw the web graph when it is much cleaner than the current web graph
is much better than the other engines at determining if a link is a true editorial citation or an artificial link
looks for natural link growth over time
heavily biases search results toward informational resources
trusts old sites way too much
a page on a site or subdomain of a site with significant age or link related trust can rank much better than it should, even with no external citations
they have aggressive duplicate content filters that filter out many pages with similar content
if a page is obviously focused on a term they may filter the document out for that term. on page variation and link anchor text variation are important. a page with a single reference or a few references of a modifier will frequently outrank pages that are heavily focused on a search phrase containing that modifier
crawl depth determined not only by link quantity, but also link quality. Excessive low quality links may make your site less likely to be crawled deep or even included in the index.
things like cheesy off topic reciprocal links are generally ineffective in Google when you consider the associated opportunity cost
Ask
looks at topical communities
due to their heavy emphasis on topical communities they are slow to rank sites until they are heavily cited from within their topical community
due to their limited market share they probably are not worth paying much attention to unless you are in a vertical where they have a strong brand that drives significant search traffic
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